According to the soil specialization (And some botanical science…)

Chamaedaphne calyculata

Ericaceae family:
Andromeda sp.
Calluna sp.

Ericaceae family:
Vaccinium…

Chamaedaphne calyculata

Plants of the olygotrophic swamps

An olygotrophic swamp

Khrugloe swamp in the Arkhangelsk region
It’s surface is covered with Sphagnum moss!

There are a lot of ecological factors, which is Chamaedaphne affected by.
I

Ecological factors
Lack of Nitrogen
Moisture
Drought
Low temperature
Excess of sun rays

Special characteristics
Ecological factors

Due to such severe conditions, plants of swamps, or OXYLOPHYTES as Chamaedaphne, have some special characteristics.

The leaves
1. The leaves
Drought

Drought

Drought
Excess of sun rays

leaves of Andromeda
are placed at an angle to the horizon.
Excess of sun rays

2. The stem
FELLOGENE cells
FELLEM, or CORK
Lack of Nitrogen

The stem
Stem of Chamaedaphne, and FELLOGENE cells
The stem
Lack of Nitrogen

The stem. Tannins in the cork
A stem of Andromeda, Ericaceae

….has some special cells, which produce fellem.
…имеют специальные клетки, производящие пробку.

1. Lack of nutrition
Olygotrophic swamps are water reservoirs which are poor with organic matter, actually with nitrogen compounds.

For example, plants of Ericaceae family –
Например, растения семейства Вересковых -

- Andromeda polifolia,
- Подбел многолистный,

- Chamaedaphne calyculata, …
…Мирт болотный,…

These cells, or FELLOGENE cells, also are A BORDER between a stem and some outer tissues, which are to be separated from the stem.
Эти клетки, или клетки ФЕЛЛОГЕНА, также находятся на ГРАНИЦЕ стебля и других тканей коры, которые затем отваливаются от стебля.

So the parts of tissues located ABOVE the FELLOGENE separate from the stem. It leads to decrease in nutrition losses during growing season.

2. Low temperature
These swamps are regions with relatively low-temperatured soil.
Олиготрофные болота – зоны со сравнительно низкой температурой почвы.

It influences badly on roots, because absorption of water is complicated.
Это плохо влияет на корни, т.к. всасывание воды [при низких температурах] затруднено.

On the other side, the sun rays reach plants very fast, which makes all the aerial runners warmer.

But leaves of Chamaedaphne (мирт) also have double layers of photosynthetic tissue to accumulate more sun rays.

As a result, subterranean organs of OXYLOPHYTES are subjected to cold, but aerial ones are overheated.

Tannins in the cork
A stem of Andromeda

It contributes to decrease influence of solar rays.

3. Water excess
Finally, the soil of olygotrophic swamps is very MOISTENED, so athmosphere OXYGEN can not penetrate through it.

3. The roots
H+
H+
H+
H+
H+
H+
H20
H20
H20
H20
Moisture

According to this, roots do not get enough oxygen for respiration.

And OXYLOPHYTES have some adaptations for breath:
They possess special tissue termed AERENCHIMA. It contains cells and plenty of air cameras!

Aerochima in
Menyanthes trifoliata rhizome
(вахта трехлистная)

Aerochima of Eriophorum (пушица):
stem
leave

There are much more ecological factors, which influence on the OXYLOPHYTES; these factors caused evolutionary adaptations in plants.

So the plants of olygotrofic swamps are adapted to high and low temperatures, lack of nutrients and oxygen.